Table 3. Dental character distribution in the Homo groups and status of H. floresiensis.
EP | MP | LP | Status a | |||||||
---|---|---|---|---|---|---|---|---|---|---|
Character | Refer to | H. hab. | H. erg. | Dmanisi | eJHe | E-Asia | H. sap. | H. flo. | H. flo. | |
#1 | C1 distal shoulder low (vs. high) | 1 in Table 2, Fig 7A | yes | − | − | yes | yes | no | yes | EP-MP |
#2 | P3 & P4 transverse crest often present | 2 & 3 in Table 2 | yes | yes? | ? | yes | no? | no | yes | EP |
#3 | P3 buccal groove(s) always present | 4 in Table 2 | yes | no | no | no | no | no | no | post-Hh |
#4 | P4 crown buccal ≤ lingual | 6 in Table 2 | yes | ? | yes | yes | no | no | yes | EP |
#5 | P3 crown long MD & asymmetric | PCs1 & 2 in Fig 4A | yes | yes | − | yes | − | no | yes | EP |
#6 | P3 lingual cusp located mesially (vs. distally on occasion) | 7 in Table 2 | yes | no | no | yes | no | no | no | post-Hh |
#7 | P3 mesiolingual crown normal (vs. beveled and wrinkled) | 9 in Table 2 | yes | yes | yes | yes | yes | yes | no | unique |
#8 | P3 buccal basal enamel swelling | 14 in Table 2 | yes | no | yes | yes | yes | no | yes | EP-MP |
#9 | P3 root often bifurcated (vs. Tomes' form or single) | 15 in Table 2 | yes | yes | yes? | yes | no | no | yes | EP |
#10 | P4 crown oblique (vs. MD symmetric) | PC2 in Fig 4C | yes | yes | − | yes | − | variable | yes | EP range |
#11 | P4 crown triangular (vs. more circular) | PC3 in Fig 4D | yes | variable | − | no | − | variable | no | post-Hh |
#12 | P4 transverse crest absent (vs. present) | 11 in Table 2 | yes | no | no | no | no | often yes | no | post-Hh |
#13 | M1 crown long & parallelogram (vs. short & rectangle) | PCs1 & 2 in Fig 5A | yes | yes | − | no | − | variable | no+ | unique |
#14 | M1 crown distobuccal corner projected (vs. abbreviated) | PC3 in Fig 5B | yes | yes | − | yes | − | variable | yes | EP range |
#15 | M2 crown long MD and distally tapering | PCs1 & 4 in Fig 5C and 5D | yes | yes | − | yes | − | variable | yes | EP range |
#16 | M2 crown parallelogram (vs. inverted trapezoid) | PC2 in Fig 5C | yes | yes | − | no | − | variable | no | post-Hh |
#17 | M1 crown long MD (vs. short) | PC1 in Figs 6A and 7C | yes | yes | − | no | − | no | no+ | unique |
#18 | M1 crown squarish (vs. pentagonal) | PC3 in Fig 6B | yes | variable | − | yes | − | variable | yes | EP range |
#19 | M1 five-cusped (vs. four-cusped) | 17 in Table 2 | yes | yes | yes | yes | yes | not always | no | Hs |
#20 | M2 crown long MD (vs. short) | PC1 in Fig 6C | yes | variable | − | no | − | variable | no | post-Hh |
#21 | M2 crown rounded (vs. square) | PC3 in Fig 6D | yes | yes | − | yes | − | variable | yes | EP range |
#22 | M2 five-cusped (vs. four-cusped or less) | 18 in Table 2 | yes | yes | yes | no? | yes | no | no | Hs? |
#23 | M1 mid-trigonid crest absent or rare | 19 in Table 2 | yes | ? | no | no | ? | yes | no | post-Hh |
#24 | Relative P3 size | 21 in Table 2 | moderate | moderate | moderate | moderate | moderate | small | large | unique |
#25 | M1/M2 size proportion | 22 in Table 2, Fig 7D | M1 < M2 | M1 ≤ M2 | M1 ≤ M2 | M1 < M2 | M1 = M2 | M1 ≥ M2 | M1 = M2 | post-Hh |
#26 | Alveolar arcade index (length-breadth index [%]) | 23 in Table 2 | narrow | wide? | narrow | moderate | wide | − | moderate | eJHe |
EP = Early Pleistocene; MP = Middle Pleistocene; LP = Late Pleistocene; H. hab. = H. habilis; H. erg. = H. ergaster; eJHe = early Javanese H. erectus; MP E-Asia = East Asian Middle Pleistocene archaic Homo; H. sap., Hs = H. sapiens; H. flo. = H. floresiensis.
aStatus of H. floresiensis is categorized as follows: EP = a primitive condition shared with the Early Pleistocene Homo; EP-MP = a primitive condition shared with the Early-Middle Pleistocene Homo; EP range = within the range of the Early Pleistocene Homo, which is mostly encompassed by the large variation of H. sapiens; post-Hh = derived from the H. habilis condition; Hs = derived like H. sapiens; unique = unique among all the samples compared; eJHe = similar to the condition of early Javanese H. erectus.