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. 2015 Nov 25;2(11):150486. doi: 10.1098/rsos.150486

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© 2015 The Authors. Published by the Royal Society under the terms of the Creative Commons Attribution License http://creativecommons.org/licenses/by/4.0/, which permits unrestricted use, provided the original author and source are credited.

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Figure 2. — The two patterning strategies. (a) In the direct strategy, target genes are controlled directly by averaged morphogen $\hat{c}$ with only one ‘access’ operation. (b) The two-tier strategy involves a second patterning factor ${\hat{c}}^{(λ)}$ ; target genes are separated from the input by two tiers of ‘access’ operations. I_raw, raw information content of the controlling morphogen; I_acc, accessible information.