Clinal variation in survival and fecundity components of fitness in two Colorado gardens. We included genotype as a random effect in these models. The low elevation CO garden included two cohorts of experimental transplants from the same 24 maternal families, whereas there was only one cohort planted into the high elevation garden. For that reason, we could not model garden by cohort interactions. Family means for the high elevation CO garden excluded plants that died because of gopher activity for the CO 2012 high elevation garden. Family means for the low elevation CO 2011 cohort excluded plants that died over the first winter (2011-2012) because excessive mortality was driven by strong runoff from snowmelt. Including those early overwinter deaths does not fundamentally alter fitness clines (Online Appendix 6).