Demographic parameters from empirical values of a natural population of A. percula. refers to the average biannual mortality rate per capita of juveniles, and adult males and females for each subpopulation between two periods: 2009–2011 and 2011–2013 (see the electronic supplementary material, table S3 for mortality rates for each of the two 2-year periods). ni is the maximal average number of juveniles per anemone in the subpopulation i between 2007 and 2013 and Ki is the total number of anemones in subpopulation i (equal to the maximal number of males or females per subpopulation).