Mass balance model used to estimate retention of P in reservoirs. Fin,i is the influx of the ith P pool into the reservoir, Fi,out is the corresponding efflux out of the reservoir, F12 represents P fixation by primary productivity, F21 represents mineralization of POP, F13 and F31 are the sorption and desorption rates of dissolved P, and Fi,bur is the permanent burial flux of the ith particulate P pool in the reservoir’s sediments.