Table 4.

Parameters of genetic variability in aspen populations.

Population		N	N a	N e	H O	H E	UH E	F
Prisady	Mean	41	6.429	3.921	0.631	0.661	0.669	0.047
	s.e.		0.850	0.602	0.059	0.044	0.045	0.065
Voronezh	Mean	25	7.429	3.970	0.580	0.687	0.701	0.188
	s.e.		1.274	0.584	0.068	0.035	0.036	0.076
Yoshkar-Ola	Mean	13	4.643	2.738	0.522	0.567	0.590	0.109
	s.e.		0.541	0.343	0.072	0.043	0.045	0.087
Total mean	Mean	26.333	6.167	3.543	0.578	0.639	0.654	0.115
	s.e.	1.791	0.558	0.308	0.038	0.024	0.025	0.044

s.e.: standard error.

N: sample size.

N _a: mean number of alleles.

N _e: effective number of alleles.

H _O: observed heterozygosity.

H _E: expected heterozygosity.

UH _E: unbiased expected heterozygosity.

F: fixation index (intrapopulation coefficient of inbreeding).