Upregulation of TCA cycle proteins in LS12 cells. Analysis of the mitochondrial subproteome demonstrates that, in addition to proteins involved in β oxidation, seven of nine TCA proteins are upregulated in LS12 cells, suggesting increased TCA cycle activity in compensation for ETC defects. Gray, proteins not identified in proteomic and MS analysis; pink, proteins identified in proteomic and MS analysis as upregulated (*p<0.05); red, proteins validated in subsequent enzyme activity assays.