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(a) Schematic diagram of swimming locomotion assays. The nematodes were put into M9 buffer solution and their swimming locomotion was recorded. Three quantitative parameters were extracted from the threshold-segmented images: (b) beating frequency, and (c) doral and (d) ventral bending amplitudes normalized by the length of the animals. n ≥ 20. *P < 0.01; **P < 0.001; ***P < 0.0001.
