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Rule-breakers. Examples of functionally validated miRNA-target interactions from Arabidopsis thaliana. The miR164/NAC1 interaction on top is representative of the canonical ‘rules’ for plant miRNA targeting: Extensive pairing to positions 2–13 of the miRNA, with most or all mismatches occurring to the 3′ end of the miRNA. The other six target sites all violate these rules, but have nonetheless been experimentally proven to function in vivo.
