Figure 5. LRP1 knockdown increases PARP-1 activity-dependent phosphorylation of Rb and CDK2 in HRECs.
A–C, LRP1 knockdown increases CDK2 and retinoblastoma (Rb) phosphorylation. HRECs were transfected with LRP1 or control siRNA. Two days later, cells were incubated under normoxia (21% O2) or hypoxia (2% O2) for 2 hours. Cell lysates were immunoblotted with indicated antibodies. The ratios of the phosphorylated CDK2 and Rb to their total respective protein amounts were quantified by Image J (B and C). D, PARP-1 activity increased in LRP1 knockdown HRECs in response to hypoxia at 2% O2. ELISA enzymatic assay for PARP-1 was performed with PAR as a substrate of PARP-1 coated in the plate. Quantitative data were presented as a ratio of relative light unit to protein amount in cell lysates. E–G, PARP-1 inhibition decreases CDK2 and retinoblastoma (Rb) phosphorylation. HRECs were incubated with PARP-1 inhibitor PJ-34 at 3 μM and under normoxia (21% O2) or hypoxia (2% O2) for 2 hours. Cell lysates were immunoblotted with indicated antibodies. The ratios of the phosphorylated CDK2 and Rb to their total respective protein amounts were quantified by Image J (F and G). H–J, PJ-34 blocks the increases in CDK2 and retinoblastoma (Rb) phosphorylation following LRP1 knockdown. HRECs were transfected with LRP1 or control siRNA. Two days later, cells were treated with PARP-1 inhibitor PJ-34 at 3 μM and incubated under normoxia (21% O2) or hypoxia (2% O2) for 2 hours. Cell lysates were immunoblotted with indicated antibodies. The ratios of the phosphorylated CDK2 and Rb to their total respective protein amounts were quantified by Image J (I and J). K, Schematic illustration to shown how LRP1 regulates cell cycle progression, endothelial proliferation and retinal angiogenesis induced by hypoxia. *, P<0.05, compared to same cells under normoxia. #, P<0.05, compared to control HRECs exposed to same oxygen level. **, P<0.05, compared to LRP1 siRNA-transfected cells without treatment of PJ-34 inhibitors. n=3. All data were analyzed with two-way ANOVA analysis followed by the Fisher’s LSD or Turkey multiple comparison test.