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Map of study sites. Offshore (Abu Roma and Shib Nazar) and nearshore (Al Quad and Inner Fsar) coral reef sites were sampled from exposed (i.e. ocean facing) and sheltered (i.e. land facing) habitats (indicated by stars) on two sampling occasions (i.e. summer and winter). Replicate numbers are shown for each sampling event and site. Open symbols: offshore, closed symbols: nearshore, circles: exposed reef sites, squares: sheltered reef sites; grey: summer, black: winter.
