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Schematic of Mxr1p-mediated trans-activation of pACS1 in P. pastoris cells cultured in YPA. A point mutation in MXRE1 or MXRE2 abrogates Mxr1p binding to pACS1 in vitro and trans-activation of pACS1-lacZ in vivo. An increase in the distance between the two MXREs also abolishes trans-activation in vivo. Therefore, the two MXREs separated by a 30-bp region functions as an MxRU.
