. 2016 Feb 28;115(4):718–729. doi: 10.1017/S0007114515004961
© The Authors 2016
This is an Open Access article, distributed under the terms of the Creative Commons Attribution licence (http://creativecommons.org/licenses/by/4.0/), which permits unrestricted re-use, distribution, and reproduction in any medium, provided the original work is properly cited.