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. 2016 Mar 1;11(3):e0150160. doi: 10.1371/journal.pone.0150160

Table 1. Equations for the Delta method and for the two closed-population models.

Delta Method Var(Ntot)=Ntot2(var NN2/+1θnθ)
Schnabel’s Binomial N=Σi=2tCiMiΣi=2tRi+1
Schumacher & Eschmeyer N=ΣitCiMi2Σi=2tRiMi

Ntot = estimated total population present during the sample period (including grade C sharks), N = estimated recognisable population, θ = proportion of recognisable individuals in population, n = total number of individuals captured during the sample period, Ci = number of potentially recognisable individuals captured at occasion i, Mi = number of previously sighted, recognisable (‘marked’) individuals in the population immediately before sample i was taken, Ri = the number of re-sighted individuals in sample i, and t = the number of sample occasions in the sample period.