Fig. 4.

Phylogenetic relationships and divergence dates for Euarchonta and outgroups were based on TimeTree (Hedges et al. 2006; accessed June 2015) and published estimates (Prideaux and Warbuton 2010; Roberts et al. 2011; Fabrae et al. 2012; Song et al. 2012). Branch colors correspond with the presence and spectral tuning of opsin photopigments. Pseudogenization events are marked with a diagonally bisected circle. The inferred shift from UV to blue sensitivity in the SWS1 opsin of the ancestral euarchontan is marked with an arrow, along with the amino acids proposed to be responsible. Dashed branches indicate opsin polymorphism. The geological time scale is abbreviated as Pa, Paleocene; Eoc, Eocene; Oli, Oligocene; Mio, Miocene; P, Plio/Pleistocene. Treeshrew art © The Sabah Society, reproduced with permission.