Innovative PSP during winter in relation to two environmental factors in the preceding natal year, and how these varied with respect to natal year: (a) tesselated territory size, inversely related to local population density, and (b) the number of large oak trees within 50 m of each nest. In some winters, innovativeness was lower among juveniles who had been born in high local density areas, or where there were fewer oak trees. Graph based on raw data. Sample sizes shown in bars. Dark bars, number of task-solvers; white, numbers of non-solvers.