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A: Ussing's original chamber. By short-circuiting the transepithelial electrical potential, Ussing could simultaneously measure Na24 flux and current across a given sample tissue (96). This allowed him to accurately assess active transport across epithelia. I, microammeter; V, voltmeter; A, apical side of tissue; B, basolateral side of tissue. B: Koefoed-Johnsen and Ussing (48) proposed passive Na+ across the outer membrane and active exchange of Na+ for K+ at the inner membrane.
