(A) RNAP trailing-edge position (left; slope ~0)
and RNAP leading-edge position (right; slope ~1) as a function of mean
TSS for each of the 64 discriminator sequences (~16,000 templates
analyzed for each discriminator).
(B) Interpretation: changes in TSS selection result from
changes in DNA scrunching. Gray, RNAP; yellow, σ; blue, -10 element
nucleotides; purple, discriminator nucleotides; i and i+1, NTP binding sites;
red, Bpa and nucleotide crosslinked to Bpa; boxes, DNA nucleotides
(nontemplate-strand nucleotides above template-strand nucleotides; nucleotides
downstream of -10 element numbered). Scrunching is indicated by bulged-out
nucleotides. Anti-scrunching is indicated by a “stretched”
nucleotide-nucleotide linkage.