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Three hypotheses for the mechanism of subtraction. A: untuned inhibition leads to subtraction if excitatory f-I curves are linear. B: withdrawal of tuned PV inhibition leads to sharpening of subthreshold input and approximate subtraction if excitatory f-I curves are nonlinear. C: nonlinear dendritic integration allows untuned SOM inhibition to sharpen tuned excitatory input, leading to subtraction if excitatory f-I curves are nonlinear.
