a, Positive correlations with noun manipulablity ratings in Experiment 1. b–e, Other studies that implicate the similar regions (the left IPL and posterior ITS, highlighted with white circles) in action performance and action semantics. b, Activation for tools > animal words/pictures in a semantic decision task (adapted from Noppeney et al., 2006). c, Activation for literal action sentences relative to those with abstract verbs (adapted from Desai et al. (2013)). d, Activation for physically using tools relative to moving a bar-like object (adapted from Brandi et al. (2014)). e, Lesion map of areas associated with producing grasping errors in patients relative to those patients who do not exhibit grasping errors (adapted from Randerath et al. (2010)). The color scale applies only to a.