Outer Plexiform Layer |
σC
|
0.05° |
Centre gaussian’s sigma |
τC
|
10 ms |
Centre signal low pass filtering time constant. |
τU
|
100 ms |
Undershoot high pass filtering time constant. |
wU
|
0.8 |
Undershoot transient relative weight. |
σS
|
0.15° |
Surround gaussian’s sigma |
τS
|
4 ms |
Surround signal low pass filtering time constant. |
λOPL
|
10 Hz/Lum. unit |
Overall gain of the centre-surround filter. |
wOPL
|
1 |
Relative weight of centre and surround signal. |
Use leaky heat equation |
True |
Averaging by gap junctions rather than dendritic spread. Leads to a non-separable spatio-temporal filter, but somewhat more realistic. |
Bipolar Cells |
λOPL’
|
50 |
Another gain applied right after λOPL, thus without biological meaning, but useful for implementation issues. |
gA0
|
50 Hz |
Inert leaks in membrane integration. |
σA
|
0.2° |
Size of the spatial neighbourhood used to estimate local contrast. |
τA
|
5 ms |
Size of the temporal neighbourhood used to estimate local contrast. |
λA
|
0 Hz |
Strength of the gain control feedback loop (no contrast gain control in primate midget cells) |
Inner Plexiform Layer |
τG
|
20 ms |
High pass filtering time constant. |
wG
|
0.7 |
Transient relative weight. |
σG
|
0° |
No additional pooling for midget cells. |
vG0
|
0 |
Bipolar linear threshold. |
λG
|
100 Hz |
Slope in the linear area. |
iG0
|
37 Hz |
This is below the threshold current (50 Hz). Thus in the dark the threshold is reached only because of the noise (see below), which leads to a irregular Poisson-like spontaneous activity (at ~1 Hz). |
Retinal Ganglion Cells (RGC) |
gL
|
50 Hz |
Leak conductance (thus the membrane time constant is 20 ms) |
σv
|
0.1 |
Gaussian white noise current’s normalized amplitude. Integration of this current by the RGCs leads to a Gaussian auto-correlated process with time constant 1/gL and variance σv. |
ηrefr
|
3 ms |
Refractory period |
Density |
20 cells/° |
RGC density (for each polarity). That is a mean inter-RGC interval of 0.05° |