Quantifiable Parameter	Observational Parameters
Velocity	Lethargy/Hyperactivity
Distance Moved	Loss of equilibrium
Duration of time spent in zone	Spasms/tremors/upside down
Zone transition-number of times organism moves between selected zones	Position in the water column
Heading to point-calculates the deviation of the animal’s path towards a point of interest	Respiration (fast/slow)
Heading calculates the heading of the selected body point	Coloration
Turn angle-difference in heading between two samples	Hiding
Angular velocity-calculated by dividing the turn angle by the sample interval
Meander-calculated by dividing the turn angle by the distance moved. Used to compare turning in animals moving at different speeds
Time spent moving
Mobility state-calculates the duration for which the complete area detected as animal is changing, even if the center point remains the same
Rotation-one rotation is completed when the selected body point has a cumulative turn angle of 360°. Turns in the opposite direction of less than the threshold are ignored.
Mobility continuous-calculates the percentage of mobility for the complete area of the detected animal, even if the center point remains the same.
Distance between subjects-calculates distance between all actors and the selected receivers
Proximity-calculates the duration for which the actor is or is not in proximity to the receiver
Relative Movement
Net weighted movement-The movement of the actor to (positive) and from (negative) the receiver, weighted by the distance between them
Weighted movement from-movement of the actor from the receiver, weighted by the distance between them.
Weighted movement to- movement of the actor to the receiver, weighted by the distance between them
Trial control state- period between two events of the Trial Control elements, or within one element
Trial control event-moment in which an event within an element you have defined in Trial Control occurs.