(A) Three icosahedral 2-fold axes lie in the equatorial plane normal to a 3-fold axis (out of page). (B) The unit cell structure of the proposed packing model for the rhombic crystal. (C)–(F) TYMV particles at the nearest-neighbor contact in the rhombic model. (C) A side-view schematic defining the coordinate axes. (D) Side view, (E) contact-plane view, and (F) top view of the two particles in the contact region, based on an interparticle spacing of 28.0 nm and the known TYMV structure.18 In (E), the pentagonal knobs are marked (“5”), and the hexagonal knobs are unmarked.