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Control of biofilm development in Vibrio cholerae at low cell density. At low concentration of autoinducers, histidine kinases LuxP and CpqS are phosphorylated and able to phosphorylate regulator LuxO. LuxO–P activate the expression of Qrr 1–4 RNAs, which, in turn, positively influence the level of c-di-GMP. c-di-GMP activates VpsR and VpsT proteins, which positively regulate biofilm genes
