The effects of sex linkage of N-mt genes on cotransmission, coevolution, genomic conflict, speciation, and ornamentation. Compatibility of nuclear and mt genes is color-coded: Blue is compatible with blue and orange with orange. The effects on F1 crosses are shown. The red star indicates offspring with total mt/N-mt incompatibility. Assuming that function is retained so long as one set of compatible genes is retained, then negative F1 effects occur only with Z linkage. For autosomal position and X linkage, incompatibility and function loss do not occur until the F2 hybrid generation.