Sampling locations for water vole used in this study. British numbered locations key: England: 1, Somerset (187); 2, Surrey; 3, Hertfordshire; 4, Cambridgeshire; 5, Norfolk; 6, Shropshire; 7, Staffordshire; 8, Derbyshire (104,106,107); 9, Lincolnshire Read's Island (195); 10, Yorkshire; 11, Northumberland (200). Wales: 12, Gwent (158,159). Scotland: 13, Fife; 14, Morayshire. European sampling locations are denoted by country name. Bracketed numbers refer to the individual sampling numbers of water vole that are directly referred to in the text.