Table 1.

17 behavioral parameter.

Parameters are defined in more details in the methods.

DOI: http://dx.doi.org/10.7554/eLife.11092.013

Motor parameter	What it represents	How it is calculated (see Materials and methods for details)
Attraction index	Overall time spent inside the odor zone	$\frac{\left(\text{Time spent inside in the during period}\right)-\left(\text{Time spent inside in the before period}\right)}{\left(\text{Time spent inside in the before period}\right)}$
Time spent/transit	Median time spent inside the odor zone per visit	$\frac{\left(\text{median time inside per transit 'during'}\right)-\left(\text{median time inside per transit 'before'}\right)}{\left(\text{median time inside per transit 'before'}\right)}$
Time to return	Median time spent outside the odor zone between successive entries into the odor-zone	$\frac{\left(\text{median time outside per transit 'during'}\right)-\left(\text{median time outside per transit 'before'}\right)}{\left(\text{median time outside per transit 'before'}\right)}$
Radial density	Mean distance from the center of the arena	Fly’s location as the distance from the center of the arena. Data were binned to 12 bins and normalized by area of each bin.
Speed inside	Mean speed inside the odor zone over the entire period	$\frac{(\text{average speed inside }\text{'}\text{during}\text{'})-(\text{average speed inside }\text{'}\text{before}\text{'})}{\left({\text{average speed inside 'before}}^{\text{'}}\right)}$
Speed outside	Mean speed outside the odor zone over the entire period	$\frac{(\text{average speed outside 'during}\text{'})-(\text{average speed outside 'before}\text{'})}{(\text{average speed outside 'before}\text{'})}$
Speed crossing inside	Acute change in speed in the first 3 s after entering the odor-zone	$\frac{\text{speed crossing inside }\text{'}\text{d}\text{uring}\text{'}-\text{speed crossing inside 'before}\text{'}}{\text{speed crossing inside 'before'}}$
Speed crossing outside	Acute change in speed in the first 3 s after leaving the odor-zone	$\frac{\text{speed crossing outside }\text{'}\text{d}\text{uring}\text{'}-\text{speed crossing outside }\text{'}\text{b}\text{efore}\text{'}}{\text{speed crossing outside 'before'}}$
Run duration	Average duration of runs	$\frac{\text{run duration 'during'}}{\text{run duration 'before'}}$
Stop duration	Average duration of stops	$\frac{\text{stop duration 'during'}}{\text{stop duration 'before'}}$
Run probability in	Fraction of time a fly spends running while inside the odor zone	$\frac{\text{probability that a fly is moving inside 'during'}}{\text{probability that a fly is moving inside 'before'}}$
Run probability out	Fraction of time a fly spends running when outside the odor zone	$\frac{\text{probability that a fly is moving ourside 'during'}}{\text{probability that a fly is moving outside 'before'}}$
Angular speed inside	Angular speed change inside the odor zone	$\frac{(\text{average angular speed inside }\text{'}\text{d}\text{uring}\text{'})-(\text{average angular speed inside }\text{'}\text{b}\text{efore}\text{'})}{(\text{average angular speed inside }\text{'}\text{b}\text{efore}\text{'})}$
Angular speed outside	Angular speed change outside the odor zone	$\frac{(\text{average angular speed outside }\text{'}\text{d}\text{uring}\text{'})-(\text{average angular speed outside }\text{'}\text{b}\text{efore}\text{'})}{(\text{average angular speed outside }\text{'}\text{b}\text{efore}\text{'})}$
Smooth turns in	Fraction of time a fly is performing a smooth turn inside the odor zone	$\frac{\text{Number of frames of smooth turns inside the odor zone}}{\text{Number of frames of run inside the odor zone}}$
Smooth turns out	Fraction of time a fly is performing smooth turns outside the odor zone	$\frac{\text{Number of frames of smooth turns outside the odor zone}}{\text{Number of frames of run outside the odor zone}}$
Sharp turns at boundary	Fraction of sharp turns near the odor boundary	The fraction of sharp turns which took place at the odor border, i.e. in a ring 2 mm around the border