qRT-PCR analysis revealed increases in expression of (A) CBC stem cell markers in Bves−/− enteroids compared to WT. Lgr5, Ascl2, Olfm4, Nanog, and Sox9 mRNA levels in Bves−/− enteroids compared to WT. qRT-PCR analysis also revealed increases in (B) +4 stem cell markers Bmi1 and Lrig1 mRNA levels in Bves−/− enteroids compared to WT with no significant differences in TERT expression (P=0.11). Expression of (C) Wnt ligands, (D) Wnt receptors, and (E) Wnt targets were significantly elevated in the Bves−/− enteroids. *P<0.05, **P<0.01 (n=6).