Figure 1. Striking and prey capture behaviors of the goblin shark.
(a) Striking behavior of the 2008 shark arranged in sequential order in milliseconds (ms). Resting phase, 0 ms; expansive phase, 0–146 ms; compressive phase, 146–785 ms (shooting stage, 146–239 ms; grasping stage, 239–319 ms; holding stage, 319–785 ms); recovery phase, 785–1,397 ms (re-opening stage, 785–1,077 ms; re-closing stage, 1,077–1,397 ms). (b) Lateral view of peak protrusion in the 2008 shark (279 ms in Fig. 1a), and lateral, frontal, and ventral views of peak protrusions in the 2011 shark. “Lateral” and “frontal” views are prey capture behaviors on a bait fish. (c) Skeletal elements related to predatory behavior of the goblin shark. bh, bathyhyal cartilage; cc, chondrocranium; ch, ceratohyal cartilage; hm, hyomandibular cartilage; lj, lower jaw (Meckel’s) cartilage; uj, upper jaw (palatoquadrate) cartilage. (a,b) Photographs are used with permission of NHK, NEP and the Discovery Channel).