Table 2.

Rates of turnover, diversification and fossil ‘discovery’ prior to a lineage's extinction (Myr⁻¹) inferred with FBD applied to Fagus and Osmundaceae. Means of oldest, random, and youngest as in table 1. Turnover rate r = μ/λ; diversification rate d = λ − μ; probability of fossil observation prior to species extinction s = ψ/(μ − ψ); speciation rate λ; extinction rate μ; fossil recovery rate ψ. Meaning of oldest, random, and youngest as in table 1.

	oldest	random	youngest
Fagus
r	0.8709	0.7379	0.8423
d	0.018	0.0197	0.0198
s	0.3015	0.6154	0.3792
λ	0.1394	0.0752	0.1256
μ	0.1214	0.0555	0.1058
ψ	0.0524	0.0887	0.0646
Osmundaceae
r	0.7249	0.58380	0.6856
d	8.73 × 10−3	8.30 × 10−3	7.88 × 10−3
s	0.3932	0.5831	0.3754
λ	0.0317	0.0199	0.0251
μ	0.0230	0.0116	0.0172
ψ	0.0149	0.0163	0.0103