Table 1.
The phenotype (or fitness) of offspring as a function of their genotype and the genotype of their mother
| Offspring genotype | |||||
|---|---|---|---|---|---|
| A 1 A 1 | A 1 A 2 | A 2 A 1 | A 2 A 2 | Maternal litter average | |
| Maternal genotype | |||||
| A 1 A 1 | 1 + am + ao | 1 + am + do | — | — | μ + aop 1 + dop 2 + am |
| A 1 A 2 | 1 + dm + ao | 1 + dm+ do | 1 + dm+ do | 1 + dm – ao | μ + ½do + ½ao(p 1 – p 2) + dm |
| A 2 A 2 | — | — | 1 –am+ do | 1 –am – ao | μ – aop 2 + dop 1 – am |
| Offspring average | μ + amp 1 + dmp 2 + ao | μ + amp 1 + dmp 2 + do | μ –amp 2 + dmp 1 + do | μ –amp 2 + dmp 1 – ao | |
Offspring genotypes have alleles ordered by their parent‐of‐origin, with the maternally inherited allele appearing first, whereas maternal genotypes are unordered (since the values are identical for the maternal genotype reciprocal heterozygotes). Marginal means are given for each maternal and offspring genotype for the case without inbreeding (see Table S1 for the values under inbreeding). Cells containing a “—” indicate combinations that do not exist (after Table 4 in Wolf and Cheverud 2012).