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Lateral stretching frequency (a) and duration (b) of burrowing, climbing and upright standing per day per rat at 3, 8 and 13 months of age in semi-naturalistic cages. Data are mean ± s.e. based on four cages housing five rats at 3 months, and six cages housing five (n = 4) or four (n = 2) rats at 8 and 13 months; *p < 0.05 and ***p < 0.001.
