Table 4. Model selection for all tested GEE models under both Brownian and Ornstein-Uhlenbeck modes of evolution using the complete dataset collected here (models from Table 2).

Model	Brownian Evolution QIC	O-U Evolution QIC
i) BM	186.365	37.459
ii) BM + CS	236.149	40.446
iii) BM + HP + CS	200.492	37.920
iv) BM+ MP + FP + CS	239.241	44.073
v) BM + HP + FP + MP + CS	200.102	41.879
vi) BM + HT	453.644	53.225
vii) BM + CS + HT	458.322	58.564
viii) BM + HP + CS + HT	468.529	60.486
ix) BM+ MP + FP + CS + HT	459.464	62.541
x) BM + HP + FP + MP + CS + HT	469.980	64.437
xi) BM + ME	188.630	37.866
xii) BM + CS + ME	242.280	41.656
xiii) BM + HP + CS + ME	215.036	39.304
xiv) BM+ MP + FP + CS + ME	245.455	45.204
xv) BM + HP + FP + MP + CS + ME	214.440	43.272

The model with the lowest QIC value (bold) represents the best balance of explanatory power and model complexity available from the suite of tested models. Models within 4 QIC of the best model (bold italics) still contain some explanatory power.