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. Author manuscript; available in PMC: 2017 Jul 1.
Published in final edited form as: Pharmacol Biochem Behav. 2016 May 14;146-147:60–72. doi: 10.1016/j.pbb.2016.05.006

Table 5.

Adult and juvenile manipulations associated with anxiety behaviors and epigenetic modifications

Brain region Manipulation Primary findings Findings by sex Species Reference
Males Females
Hippocampus Kap1 knockout ↓ H3K9me3 at Mkrn3 &
Pcdhβ6
-- TG mice (Jakobsson et al., 2008)
↑ H3 & H4 acetylation at
Mkrn3 & Pcdhβ6
--
High anxiety strain Dnmt1 expression in
DG & CA3
-- bLR-HR rats (Simmons et al., 2013)
Dnmt1 expression in
CA1
--
High anxiety strain Fgf2 methylation bLR-HR rats (Chaudhury et al., 2014)
↑ H3K9me3
Methyl-donor-deficient
diet
Dnmt3a & Dnmt3b
expression
-- C57BL/6J mice (Ishii et al., 2014)
Corticosterone
administration
Fkbp5 methylation -- C57BL/6J mice (Yang et al., 2012)
Dnmt1 expression --
Chronic unpredictable
stress
↓ H3K9 & H4K12
acetylation
-- SD rats (Liu et al., 2014)
Hdac5 expression --
Amygdala miR-34c
overexpression
↑ anxiolytic behaviors -- TG & C57BL/6J mice (Haramati et al., 2011)
Chronic mild stress Crhr1 methylation -- Inbred strain (CD1
background)
(Sotnikov et al., 2014)
High anxiety strain Dnmt1 expression in BL
& L
-- bLR-HR rats (Simmons et al., 2013)
Dnmt1 expression in
MN
--
High anxiety strain Fgf2 methylation bLR-HR rats (Chaudhury et al., 2014)
↑ H3K9me3
Nucleus
Accumbens
High anxiety strain ↑ H3K9me3 bLR-HR rats (Chaudhury et al., 2014)
Hypothalamus Social defeat stress Crf methylation -- C57BL/6J mice (Elliott et al., 2010)
Dnmt3b & Hdac2
expression
--
Corticosterone
administration
Fkbp5 methylation -- C57BL/6J mice (Lee et al., 2010)
Raphe Nuclei miR-135 knockdown ↑ anxiogenic behaviors -- C57BL/6 & TG mice (Issler et al., 2014)
Prefrontal
Cortex
Mll1 deletion ↓ H3K4me3 at multiple
promoter regions
TG mice (C57BL/6J
background)
(Jakovcevski et al., 2015)
Gomafu knockdown ↑ anxiolytic behaviors -- C57BL/6 mice (Spadaro et al., 2015)

✓ = Significant; -- = Not Included; SD = Sprague Dawley; TG = Transgenic; bLR-HR = Bred Low Responders and Bred High Responders.