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The mad1-RLK-AAA mutant participates in a SAC response when associated with Kre28. Serial dilutions of mad1∆ GFP strains shows that cells with a plasmid encoding mutant mad1-RLK/AAA-GBP, which normally does not complement mad1∆, rescues benomyl sensitivity in cells containing GFP-tagged Kre28, Cdc20, and Ipl1.
