Phylogenetic PCA of phylogenetic residuals of lemur morphological measurements projected into morphological space. Colored phylogenetic branches indicate time since the root. Tips of the phylogeny are abbreviated: A, Avahi; B, Babakotia; C, Cheirogaleus; D, Daubentonia; E, Eulemur; H, Hapalemur; Ha, Hadropithecus; I, Indri; L, Lemur; Le, Lepilemur; M, Megaladapis; Me, Mesopropithecus; Mc, Microcebus; Mz, Mirza; P, Propithecus; Pa, Pachylemur; Pr, Paleopropithecus; V, Varecia. Tips with colored white circles indicate extant taxa, and tips with orange circles indicate extinct taxa. Colored polygons represent seed dispersers vs. nondispersers, with darker shaded areas indicating the morphological space occupied solely by extant lemurs. Lemur skull illustrations show morphologies occupying particular areas of morphological space. MA, million years before present.