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Two different views of the overall structure of the actin–Arp4–HSA complex. Actin, orange; Arp4, green; HSA domain, magenta; the “hydrophobic plug” of actin, blue; the two insertions (In1 and In2) of Arp4, red. The N and C termini of the HSA domain are labeled with black dots. The four subdomains of actin are labeled D1, D2, D3, and D4.
