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. 2016 Aug 23;6:31857. doi: 10.1038/srep31857

Table 3. Phenotypic analysis of F2 offspring of grandparents (F0 mothers and fathers) that were fed a high fat (HFD) or control diet for 14 weeks.

  Male
Female
Control Maternal grandfather Maternal grandmother Paternal grandfather Paternal grandmother Control Maternal grandfather Maternal grandmother Paternal grandfather Paternal grandmother p
N (litters) 14 (7) 8 (4) 8 (4) 10 (5) 10 (5) 14 (7) 8 (4) 8 (4) 10 (5) 10 (5)  
Body size
Weight (g) 416 ± 5.2 448 ± 13.4 446 ± 8.8 412 ± 11.4 433 ± 9.5 268 ± 7.6 277 ± 5.4 269 ± 7.5 262 ± 5.9 271 ± 5.3 0.124
Length (cm) 23.3 ± 0.1 23.5 ± 0.2 23.5 ± 0.2 22.8 ± 0.1 23.4 ± 0.2 21.1 ± 0.2 20.9±0.1 21.0 ± 0.2 20.7 ± 0.2 21.1 ± 0.2 0.613
Organ weights
Liver/bw (mg/g) 43.2 ± 0.8 40.8 ± 0.5 42.3 ± 1.4 42.8 ± 1.2 44.4 ± 1.4 38.9 ±  ± 0.9 37.0 ± 1.5 34.3 ± 1.5 38.4 ± 1.4 37.6 ± 1.1 0.24
Pancreas/bw (mg/g) 2.6 ± 0.1 2.4 ± 0.2 2.1 ± 0.1 2.3 ± 0.1 2.2 ± 0.1 3.5 ± 0.1 3.5 ± 0.2 3.8 ± 0.4 4.0 ± 0.2 3.0 ± 0.2 0.007
Adiposity
Retroperitoneal fat/bw (mg/g) 4.8 ± 0.4 6.6 ± 0.5 5.1 ± .4 6.3 ±  ± 0.5 6.1 ± 0.4 4.1 ± 0.5 3.6 ± 0.2 3.9 ± 0.7 4.9 ± 0.3 5.2 ± 0.3 0.003
 Gonad fat/bw(mg/g) 6.6 ± 0.3 8.4 ± 0.6* 7.0 ± 0.2 7.9 ± 0.5 6.9 ± 0.4 5.9 ± 0.5 5.2 ± 0.3 5.9 ±  ± 0.4 6.3 ± 0.4 6.3 ± 0.5 0.017
Adiposity index (mg/g) 18.0 ± 1.0 23.4 ± 1.7 19.0 ± 0.7 22.0 ± 1.3 19.9 ± 1.0 16.0 ± 1.3 14.0 ± 0.6 15.7 ± 1.3 17.4 ± 0.9 17.8 ± 1.2 0.012
Biochemistry
Insulin AUC (μg/l.min) 90.2 ± 10.7 148.7 ± 18.2* 104.0 ± 7.1 119.0 ± 13.3 128.5 ± 18.6 60.3 ± 7.4 86.2 ± 16.2 48.4 ± 2.9 103.3 ± 8.4 72.0 ± 11.9 0.016
Glucose AUC (mM.min) 964 ± 18.5 904 ± 6.6 947 ± 36.5 954 ± 10.7 966 ± 6.4 947 ± 24.1 931 ± 18.9 921 ± 27.7 1146 ± 13.4 978 ± 9.1 0.15
Leptin (ng/ml) 2.5 ± 0.3 5.0 ± 0.8 3.3 ± 0.2 3.6 ± 0.5 2.9 ± 0.4 1.8 ± 0.2 1.7 ± 0.2 1.9 ± 0.2 1.6 ± 0.1 1.9 ± 0.2 0.001
Cholesterol (mM) 1.08 ± 0.03 1.16 ± 0.05 1.03 ± 0.07 1.18 ± 0.05 1.03 ± 0.04 1.02 ± 0.04 1.10 ± 0.08 1.06 ± 0.06 1.25 ± 0.06 1.08 ± 0.07 0.474
Triglycerides (mM) 1.08 ± 0.05 1.3 ± 0.189 1.54 ±  ± 0.13 1.16 ± 0.10 1.27 ± 0.12 0.82 ± 0.05 0.78 ± 0.04 1.21 ± 0.12 0.83 ± 0.07 1.14 ± 0.13 0.106
Reproduction
LH (ng/ml) 0.6 ± 0.1 1.2 ± 0.2 0.7 ± 0.7 1.0 ± 0.2 1.0 ± 0.2           0.250^
Testosterone(ng/ml) 8.5 ± 0.5 6.1 ± 0.8 8.6 ± 1.4 8.7 ± 1.4 7.80 ± 0.6           0.270^
LH:T ratio 0.07 ± 0.01 0.21 ± 0.04 0.09 ± 0.02 0.13 ± 0.04 0.12 ± 0.02           0.017^
Penis Length (mm) 12.1 ± 0.1 12.3 ± 0.2 12.5 ± 0.2 12.3 ± 0.2 12.3 ± 0.1           0.614^
Sperm count (106) 6.0±12 36.3 ± 7.5 22.4 ± 7.3 33.2 ± 8.8 23.3 ±  ± 7.7           0.629^
AGD (mm) 48.6 ± 0.7 49.5 ± 0.9 49.9 ± 1.1 47.8 ± 0.8 50.9 ± 0.6 21.3 ± 0.5 23.1±0.2 21.7 ±  ± 0.4 22.4 ± 0.4 22.7 ± 0.4 0.121

Data are derived from post-mortem dissection at 19 weeks of age. Biochemical data derives from 09.00 fasting plasma obtained during glucose tolerance testing at 17 weeks of age. Data was analysed by linear mixed model with group and sex as fixed factors and litter as a random factor with post hoc Bonferroni analysis. Comparing groups within each sex, values significantly different (p < 0.05) from control are shown in bold.

*Indicates different only with least significant difference analysis (not taking multiple testing into account). ^Indicates sex not used as a fixed factor as data only available for males.