Table 3. Mixed models testing how the exposure to oxidative stress (diquat) interacted with the dietary carotenoid treatment at the end of the experiment.
The level of each dependent variable in the sampling event precedent to the diquat exposure is included as a covariate for color and blood variables. The models describe the backward step (using the P = 0.10 threshold) previous to remove the diquat × CAR interaction (i.e. when it was non-significant; Methods).
| Dependent variable | Terms in the model | Slope | SE | F | df | P |
|---|---|---|---|---|---|---|
| Eye rings redness | Carotenoid | 24.93 | 3,147 | <0.001 | ||
| Diquat | 0.03 | 1,146 | 0.859 | |||
| Sex | 0.8 | 1,146 | 0.373 | |||
| Carotenoid × diquat | 1.38 | 3,146 | 0.252 | |||
| Sex × diquat | 3.21 | 1,146 | 0.075 | |||
| Total brightness | −0.00004 | 0.00002 | 6.25 | 1,147 | 0.014 | |
| Eye ring redness in day 48 | 0.291 | 0.064 | 20.95 | 1,147 | <0.001 | |
| Liver vitamin A | 0.0004 | 0.0003 | 1.33 | 1,147 | 0.251 | |
| Eye ring tocopherol | 0.025 | 0.012 | 4.43 | 1,146 | 0.037 | |
| Eggs during diquat experiment | −0.002 | 0.0008 | 4.44 | 1,146 | 0.037 | |
| Bill redness | Carotenoid | 17.19 | 383.2 | <0.001 | ||
| Diquat | 4.23 | 176.6 | 0.043 | |||
| Sex | 1.46 | 175.9 | 0.231 | |||
| Carotenoid × diquat | 1.03 | 374.3 | 0.382 | |||
| Sex × carotenoid | 1.34 | 369.7 | 0.269 | |||
| Total brightness | −0.0001 | 0.00003 | 23.14 | 1,143 | <0.001 | |
| Bill redness in day 48 | 0.151 | 0.056 | 7.4 | 1,134 | 0.007 | |
| Bill tocopherol | 0.048 | 0.014 | 11.7 | 1,118 | <0.001 | |
| Liver vitamin A | 0.001 | 0.0004 | 5.72 | 1,138 | 0.018 | |
| Leg redness | Carotenoid | 10.55 | 3,136 | <0.001 | ||
| Diquat | 0.23 | 1,137 | 0.631 | |||
| Sex | 3.98 | 1,138 | 0.048 | |||
| Carotenoid × diquat | 0.67 | 3,137 | 0.575 | |||
| Sex × diquat | 0.33 | 1,137 | 0.567 | |||
| Sex × carotenoid | 1.69 | 3,137 | 0.173 | |||
| Red chip | 1.018 | 0.221 | 21.28 | 128.8 | <0.001 | |
| Leg redness in day 48 | 0.594 | 0.065 | 82.46 | 1,137 | <0.001 | |
| Leg tocopherol | 2.752 | 0.605 | 20.73 | 1,137 | <0.001 | |
| Liver tocopherol | −1.469 | 0.544 | 7.28 | 1,138 | 0.008 | |
| Liver vitamin A | −0.024 | 0.015 | 2.73 | 1,135 | 0.101 | |
| Total number of eggs | −0.032 | 0.014 | 5.1 | 1,137 | 0.026 | |
| Total astaxanthin in the eye rings | Carotenoid | 51.64 | 3,146 | <0.001 | ||
| Diquat | 1.47 | 1,151 | 0.227 | |||
| Carotenoid × diquat | 3.21 | 3,147 | 0.025 | |||
| Sex | 17.4 | 1,148 | <0.001 | |||
| Sex × carotenoid | 3.21 | 3,146 | 0.025 | |||
| Tocopherol in the eye rings | 0.674 | 0.06 | 125.22 | 1,149 | <0.001 | |
| Total number of eggs | −0.004 | 0.002 | 6.54 | 1,145 | 0.012 | |
| Total papilioerythrinone in the eye rings | Carotenoid | 19.9 | 388.3 | <0.001 | ||
| Diquat | 0.55 | 180.1 | 0.46 | |||
| Sex | 13.83 | 179.6 | <0.001 | |||
| Carotenoid × diquat | 0.33 | 377.4 | 0.804 | |||
| Sex × diquat | 1.74 | 181.8 | 0.19 | |||
| Sex × carotenoid | 3.71 | 379.5 | 0.015 | |||
| Fat retinol | 0.032 | 0.031 | 1.04 | 1,136 | 0.309 | |
| Plasma tocopherol | 0.3 | 0.182 | 2.71 | 1,140 | 0.102 | |
| Tocopherol in the eye rings | 0.85 | 0.149 | 32.42 | 1,140 | <0.001 | |
| Total number of eggs | −0.007 | 0.004 | 3.03 | 183.8 | 0.086 | |
| Tocopherol in the eye rings | Carotenoid | 3.64 | 371.7 | 0.017 | ||
| Diquat | 1.22 | 175.7 | 0.272 | |||
| Carotenoid × diquat | 2.63 | 373.5 | 0.056 | |||
| Total number of eggs | −0.006 | 0.002 | 8.96 | 171.1 | 0.004 | |
| Total astaxanthin in the bill | Carotenoid | 141.3 | 3,151 | <0.001 | ||
| Sex | 5.43 | 1,155 | 0.021 | |||
| Diquat | 4.68 | 1,157 | 0.032 | |||
| Carrotenoid × diquat | 2.67 | 3,155 | 0.049 | |||
| Plasma tocopherol | 1.176 | 0.061 | 371.2 | 1,158 | <0.001 | |
| Total number of eggs | −0.007 | 0.002 | 17.86 | 1,151 | <0.001 | |
| Total papilioerythrinone in the bill | Carotenoid | 134.5 | 364.8 | <0.001 | ||
| Diquat | 0.08 | 168.8 | 0.774 | |||
| Sex | 2.66 | 175.8 | 0.107 | |||
| Carotenoid × diquat | 1.76 | 366 | 0.163 | |||
| Tocopherol in the bill | 1.536 | 0.133 | 133.35 | 1,134 | <0.001 | |
| Plasma retinol | −9.373 | 4.689 | 4 | 1,140 | 0.048 | |
| Total number of eggs | −0.009 | 0.003 | 9.17 | 173.4 | 0.003 | |
| Tocopherol in the bill | Carotenoid | 2.94 | 3,158 | 0.035 | ||
| Diquat | 3.91 | 1,162 | 0.05 | |||
| Carotenoid × diquat | 3.09 | 3,160 | 0.029 | |||
| Sex | 5.6 | 1,161 | 0.019 | |||
| Total number of eggs | −0.007 | 0.002 | 9.66 | 1,159 | 0.002 | |
| Total astaxanthin in the legs | Carotenoid | 7.36 | 392.1 | <0.001 | ||
| Diquat | 0.13 | 178.8 | 0.7168 | |||
| Sex | 2.98 | 186.9 | 0.088 | |||
| Carotenoid × diquat | 0.07 | 376 | 0.974 | |||
| Sex × diquat | 0.14 | 176.6 | 0.712 | |||
| Sex × carotenoid | 0.56 | 374.3 | 0.645 | |||
| Plasma tocopherol | 0.146 | 0.113 | 1.66 | 1,144 | 0.199 | |
| Liver vitamin A | 0.004 | 0.002 | 3.32 | 1,133 | 0.071 | |
| Fat retinol | −0.032 | 0.018 | 3.19 | 1,130 | 0.077 | |
| Tocopherol in the legs | 0.526 | 0.097 | 29.25 | 1,141 | <0.001 | |
| Total number of eggs | 0.003 | 0.002 | 1.37 | 187.5 | 0.244 | |
| Total papilioerythrinone in the legs | Carotenoid | 4.17 | 392.9 | 0.008 | ||
| Diquat | 0.61 | 184.9 | 0.436 | |||
| Sex | 6.93 | 191.3 | 0.01 | |||
| Carotenoid × diquat | 0.17 | 382.1 | 0.919 | |||
| Sex × diquat | 0.24 | 181.1 | 0.627 | |||
| Sex × carotenoid | 0.16 | 378.8 | 0.919 | |||
| Tocopherol in the legs | 0.867 | 0.147 | 34.64 | 1,143 | <0.001 | |
| Liver vitamin A | −0.003 | 0.004 | 0.49 | 1,144 | 0.484 | |
| Tocopherol in the legs | Carotenoid | 4.09 | 382.2 | 0.009 | ||
| Diquat | 2.4 | 183.8 | 0.125 | |||
| Carotenoid × diquat | 1.21 | 182.4 | 0.3126 | |||
| Sex | 5.98 | 189.5 | 0.016 | |||
| Total number of eggs | −0.004 | 0.002 | 4.41 | 182.1 | 0.039 | |
| Plasma lutein | Carotenoid | 151.01 | 3,149 | <0.001 | ||
| Diquat | 0.01 | 1,149 | 0.925 | |||
| Carotenoid × diquat | 2.84 | 3,149 | 0.04 | |||
| Lutein at day 48 | 0.446 | 0.062 | 51.81 | 1,149 | <0.001 | |
| Plasma tocopherol | 0.479 | 0.0326 | 215.52 | 1,149 | <0.001 | |
| Eggs during diquat experiment | −0.003 | 0.002 | 4.42 | 1,149 | 0.037 | |
| Plasma zeaxanthin | Carotenoid | 321.37 | 3,146 | <0.001 | ||
| Diquat | 0.83 | 1,146 | 0.363 | |||
| Carotenoid × diquat | 1.57 | 3,146 | 0.2 | |||
| Zeaxanthin at day 48 | 0.307 | 0.07 | 19.24 | 1,146 | <0.001 | |
| Plasma tocopherol | 0.572 | 0.039 | 219.4 | 1,146 | <0.001 | |
| Fat tocopherol | −0.037 | 0.013 | 8.24 | 1,146 | 0.005 | |
| Liver vitamin A | 0.002 | 0.001 | 4.79 | 1,146 | 0.03 | |
| Eggs during diquat experiment | −0.004 | 0.002 | 4.05 | 1,146 | 0.046 | |
| Plasma tocopherol | Carotenoid | 5.26 | 395.1 | 0.002 | ||
| Diquat | 2.72 | 182.3 | 0.103 | |||
| Sex | 0.14 | 186.2 | 0.71 | |||
| Carotenoid × diquat | 0.7 | 379.3 | 0.552 | |||
| Sex × diquat | 0.7 | 182.7 | 0.404 | |||
| Tocopherol at day 48 | 0.199 | 0.069 | 8.31 | 1,139 | 0.005 | |
| Liver vitamin A | −0.004 | 0.002 | 3.99 | 1,134 | 0.048 | |
| Fat retinol | 0.03 | 0.016 | 3.34 | 1,138 | 0.07 | |
| Plasma retinol | 0.259 | 0.162 | 2.57 | 1,141 | 0.111 | |
| Total number of eggs | −0.003 | 0.002 | 2.21 | 184.2 | 0.141 | |
| Plasma retinol | Carotenoid | 1.36 | 380.5 | 0.262 | ||
| Diquat | 1.54 | 180.3 | 0.218 | |||
| Sex | 0.51 | 177.9 | 0.476 | |||
| Carotenoid × diquat | 0.51 | 377.7 | 0.678 | |||
| Sex × diquat | 0.31 | 178.7 | 0.579 | |||
| Sex × carotenoid | 0.64 | 379.5 | 0.589 | |||
| Retinol at day 48 | 33.86 | 4.27 | 62.89 | 1,134 | <0.001 | |
| Plasma tocopherol | 5.209 | 1.95 | 7.13 | 1,140 | 0.009 | |
| Eggs during diquat experiment | −0.258 | 0.095 | 7.34 | 173.6 | 0.008 | |
| Liver lutein | Carotenoid | 128.63 | 3,157 | <0.001 | ||
| Diquat | 0.06 | 1,157 | 0.811 | |||
| Carotenoid × diquat | 1.49 | 3,157 | 0.22 | |||
| Plasma tocopherol | 0.075 | 0.036 | 4.45 | 1,157 | 0.037 | |
| Liver tocopherol | 0.263 | 0.028 | 85.5 | 1,157 | <0.001 | |
| Liver zeaxanthin | Carotenoid | 315.42 | 3,151 | <0.001 | ||
| Diquat | 0 | 1,154 | 0.971 | |||
| Carotenoid × diquat | 3.06 | 3,151 | 0.03 | |||
| Plasma tocopherol | 0.1 | 0.046 | 4.69 | 1,151 | 0.031 | |
| Liver tocopherol | 0.341 | 0.04 | 74.08 | 140.3 | <0.001 | |
| Plasma retinol | 0.003 | 0.001 | 3.6 | 1,153 | 0.06 | |
| Liver tocopherol | Carotenoid | 12.77 | 3,161 | <0.001 | ||
| Diquat | 6.47 | 1,161 | 0.012 | |||
| Carotenoid × diquat | 2.76 | 3,161 | 0.044 | |||
| Liver vitamin A | Carotenoid | 57.35 | 3,152 | <0.001 | ||
| Diquat | 0.04 | 1,154 | 0.834 | |||
| Sex | 22.3 | 1,154 | <0.001 | |||
| Carotenoid × diquat | 0.47 | 3,152 | 0.707 | |||
| Sex × diquat | 0.71 | 1,152 | 0.399 | |||
| Plasma tocopherol | −11.032 | 4.001 | 7.6 | 1,153 | 0.007 | |
| Liver tocopherol | 10.309 | 3.538 | 8.49 | 155.8 | 0.005 | |
| Total number of eggs | −0.394 | 0.07 | 31.26 | 1,152 | <0.001 | |
| Fat lutein | Carotenoid | 12.87 | 3,147 | <0.001 | ||
| Diquat | 0.06 | 1,148 | 0.808 | |||
| Sex | 0.13 | 1,147 | 0.716 | |||
| Carotenoid × diquat | 0.21 | 3,148 | 0.89 | |||
| Sex × diquat | 0.19 | 1,147 | 0.662 | |||
| Sex × carotenoid | 1.26 | 3,147 | 0.292 | |||
| Fat tocopherol | 1.288 | 0.181 | 50.69 | 1,142 | <0.001 | |
| Liver vitamin A | 0.022 | 0.01 | 4.62 | 1,147 | 0.033 | |
| Plasma retinol | −0.039 | 0.016 | 5.77 | 1,147 | 0.018 | |
| Fat retinol | 0.241 | 0.087 | 7.72 | 1,148 | 0.006 | |
| Fat zeaxanthin | Carotenoid | 46.74 | 3,148 | <0.001 | ||
| Diquat | 0.16 | 1,148 | 0.687 | |||
| Sex | 0.28 | 1,147 | 0.598 | |||
| Carotenoid × diquat | 0.12 | 3,148 | 0.948 | |||
| Sex × diquat | 0.88 | 1,148 | 0.349 | |||
| Sex × carotenoid | 1.23 | 3,148 | 0.302 | |||
| Fat tocopherol | 0.911 | 0.15 | 37.07 | 1,129 | <0.001 | |
| Liver vitamin A | 0.019 | 0.01 | 4.55 | 1,148 | 0.035 | |
| Plasma retinol | −0.023 | 0.014 | 3.01 | 1,148 | 0.085 | |
| Fat retinol | 0.203 | 0.073 | 7.84 | 1,148 | 0.006 | |
| Fat tocopherol | Carotenoid | 0.57 | 395.7 | 0.638 | ||
| Diquat | 1.3 | 182.8 | 0.257 | |||
| Sex | 0.04 | 181.4 | 0.849 | |||
| Carotenoid × diquat | 0.15 | 382 | 0.931 | |||
| Sex × diquat | 0.31 | 181 | 0.578 | |||
| Sex × carotenoid | 1.71 | 379.4 | 0.171 | |||
| Liver vitamin A | 0.005 | 0.004 | 1.11 | 1,130 | 0.295 | |
| Plasma retinol | −0.012 | 0.007 | 3.05 | 1,138 | 0.083 | |
| Total number of eggs | −0.005 | 0.005 | 0.99 | 191.4 | 0.323 | |
| Fat retinol | Carotenoid | 29.11 | 3,149 | <0.001 | ||
| Diquat | 0.18 | 1,148 | 0.671 | |||
| Sex | 0.14 | 1,147 | 0.706 | |||
| Carotenoid × diquat | 0.24 | 3,149 | 0.867 | |||
| Sex × diquat | 0.05 | 1,149 | 0.816 | |||
| Sex × carotenoid | 1.06 | 3,148 | 0.368 | |||
| Plasma tocopherol | 0.64 | 0.535 | 1.43 | 1,149 | 0.234 | |
| Liver tocopherol | 0.136 | 0.428 | 0.1 | 1,149 | 0.752 | |
| Fat tocopherol | −0.103 | 0.165 | 0.39 | 164.6 | 0.537 | |
| Total number of eggs | −0.025 | 0.01 | 6.77 | 1,148 | 0.01 | |
| UA & ALB-corrected PLAOX | Carotenoid | 3.4 | 379.9 | 0.022 | ||
| Diquat | 0.37 | 171.4 | 0.543 | |||
| Sex | 1.61 | 369.7 | 0.209 | |||
| Carotenoid × diquat | 1.34 | 169.6 | 0.269 | |||
| Carotenoid × sex | 0.33 | 362.6 | 0.805 | |||
| Diquat × sex | 0.03 | 175.9 | 0.855 | |||
| Carotenoid × diquat × sex | 2.85 | 361.3 | 0.045 | |||
| AOX at day 48 | 0.331 | 0.11 | 9.07 | 173.1 | 0.004 | |
| Fat tocopherol | 0.054 | 0.03 | 3.21 | 1,103 | 0.076 | |
| Uric acid | 0.056 | 0.005 | 119.03 | 1,101 | <0.001 | |
| Albumin | −0.009 | 0.005 | 3.73 | 1,104 | 0.056 | |
| Liver vitamin A | −0.004 | 0.002 | 3.69 | 187.2 | 0.058 | |
| Eggs during diquat experiment | −0.011 | 0.005 | 4.66 | 183.9 | 0.034 | |
| Plasma TRG-corrected MDA | Carotenoid | 0.29 | 3,139 | 0.836 | ||
| Diquat | 6.84 | 1,139 | 0.009 | |||
| Sex | 4.8 | 1,140 | 0.03 | |||
| Carotenoid × diquat | 0.86 | 3,139 | 0.466 | |||
| Diquat × sex | 4.45 | 1,140 | 0.037 | |||
| TRG-corrected MDA at day 48 | 2.419 | 0.745 | 10.48 | 1,139 | 0.002 | |
| Triglycerides | 0.004 | 0.001 | 36.4 | 1,140 | <0.001 | |
| Eggs during diquat experiment | 0.126 | 0.044 | 8.28 | 1,140 | 0.005 | |
| Liver MDA | Carotenoid | 1.27 | 388 | 0.289 | ||
| Diquat | 0.2 | 177.4 | 0.659 | |||
| Sex | 22.76 | 180.4 | <0.001 | |||
| Carotenoid × diquat | 1.96 | 376.7 | 0.127 | |||
| Carotenoid × sex | 1.21 | 375 | 0.311 | |||
| Diquat × sex | 0.15 | 175.4 | 0.699 | |||
| Carotenoid × diquat × sex | 4.65 | 376 | 0.005 | |||
| Liver vitamin A | 0.0002 | 0.0001 | 7.07 | 1,138 | 0.009 | |
| Plasma tocopherol | 0.0003 | 0.0001 | 3.73 | 1,139 | 0.055 | |
| Heart MDA | Carotenoid | 0.09 | 3,157 | 0.963 | ||
| Diquat | 0.95 | 1,157 | 0.331 | |||
| Sex | 2.75 | 1,157 | 0.099 | |||
| Carotenoid × diquat | 1.79 | 3,157 | 0.151 | |||
| Erythrocyte resistance to oxidative stress | Carotenoid | 0.35 | 361.1 | 0.793 | ||
| Diquat | 5.66 | 161.1 | 0.021 | |||
| Carotenoid × diquat | 2.27 | 361.4 | 0.09 | |||
| Lag time | −0.229 | 0.035 | 44.06 | 1,121 | <0.001 |
Note:
ALB, albumin; MDA, malondyaldehydes; PLAOX, plasma antioxidants; TRG, tryglycerides; UA, uric acid.