Fig. 4.

Parameterization of the generative model. Untuned saccade- and fixation-related temporal responses were modeled using linear combinations of raised-cosine temporal basis functions. Spatiotemporal tuning to saliency, relevance, energy, and saccade direction were modeled using bilinear models with left multipliers representing temporal basis function loadings, and right multipliers representing spatial basis function (cosine and sine) loadings. Additionally, a spike-history (self) term was modeled using a linear combination of temporal basis functions causally aligned to spike events. Parameters of the model were fit using maximum-likelihood with elastic net regularization.