Figure 2. How morphological measurements were taken using photographs (panels A, B and C) and on board the survey (panel D).

Morphological traits were calculated from these measurements using the formulae in Table 2. (A) TL, total length; HL, head length; ED, eye diameter; SAT, surface area of tail; TH, tail height. (B) EP, eye position. (C) SAM, surface area of mouth protrusion; MA, mouth angle. (D) MW, mouth width; MH, mouth height.