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Movement and coordination were tested by placing the pups head down on a 30° inclined plane and scoring their natural response to turn themselves upright. The percentage of pups able to turn themselves upright in reaction to negative geotaxis was greatly enhanced in Pip6a-PMO–treated pups over PMO-treated and untreated pups (*P ≤ 0.01; **P ≤ 0.001; ***P ≤ 0.0001 by binomial test).
