S |
Number of synonymous sites |
N |
Number of nonsynonymous sites |
Ks |
Synonymous substitution rate |
Ka |
Nonsynonymous substitution rate |
ω |
Estimator of selective pressure, ω=Ka/Ks |
Sd
|
Number of synonymous substitutions |
Nd
|
Number of nonsynonymous substitutions |
t |
Divergence time between two sequences |
a |
The shape parameter of gamma distribution |
α |
Transitional rate |
α1
|
Transitional rate between purines |
α2
|
Transitional rate between pyrimidines |
β |
Transversional rate |
κ |
Ratio of transitional rate/transversional rate |
κR
|
Ratio of transitional rate between purines to transversional rate, κR=α1/β
|
κY
|
Ratio of transitional rate between pyrimidines to transversional rate, κY=α2/β
|
gN
|
Frequency of nucleotide N, N∈[T, C, A, G] |
gR
|
gR = gA + gG
|
gY
|
gY = gT + gC
|