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. 2009 Nov 25;7(3):116–127. doi: 10.1016/S1672-0229(08)60040-6

Table 1.

Symbols used in this paper

Symbol Description
S Number of synonymous sites
N Number of nonsynonymous sites
Ks Synonymous substitution rate
Ka Nonsynonymous substitution rate
ω Estimator of selective pressure, ω=Ka/Ks
Sd Number of synonymous substitutions
Nd Number of nonsynonymous substitutions
t Divergence time between two sequences
a The shape parameter of gamma distribution
α Transitional rate
α1 Transitional rate between purines
α2 Transitional rate between pyrimidines
β Transversional rate
κ Ratio of transitional rate/transversional rate
κR Ratio of transitional rate between purines to transversional rate, κR=α1/β
κY Ratio of transitional rate between pyrimidines to transversional rate, κY=α2/β
gN Frequency of nucleotide N, N∈[T, C, A, G]
gR gR = gA + gG
gY gY = gT + gC