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(A) Vimentin monomer structure with the central α-helical rod domain (green boxes; light and dark green for facilitating understanding of the assembly hierarchy) and intrinsically unstructured head and tail domains (black lines). Monomers form antiparallel tetramers, consisting of two coiled-coil dimers, in low-salt buffers. (B) Lateral assembly step. p tetramers associate to form a ULF. (C) Elongation reaction. Association of ULFs or filaments of any length are included in our model.
