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. 2016 Oct 14;11(10):e0164247. doi: 10.1371/journal.pone.0164247

Table 1. Motility parameters by quadrant.

Quadrant 1:Velocity is high Meandering Index is low “Active migration, returns to origin” Quadrant 2: Velocity is high Meandering Index is high “Directional, sustained motility”
Quadrant 3: Velocity is low Meandering Index is low “Low motility” Quadrant 4: Velocity is low Meandering Index is high “Non-sustained motility”