Table 4.

Results of GLMMs fitted to the numbers of Varroa mites per approx. 500 bees as well as of a logistical model applied to the presence/absence data of Nosema, DWV, SBV, and ABPV

	Varroa infestation level as determined by the icing sugar method	Nosema	DWV	SBV	ABPV
Intercept	3.48 ± 0.09***	−3.90 ± 1.00***	−3.21 ± 2.59	−45.06 ± 13,448.37	−7.71 ± 7.84
DAP	4.04 ± 3.53	−8.01 ± 10.86	0.02 ± 0.05	−6.52 ± 3,544.90	34.75 ± 42.91
Study site (test compared to reference)	0.03 ± 0.11	−1.45 ± 0.95	0.12 ± 0.39	11.43 ± 9,508.04	0.23 ± 0.48
Distance to OSR (distant compared to edge)	−0.02 ± 0.11	1.64 ± 1.04	0.41 ± 0.39	−19.56 ± 8,816.63	−0.44 ± 0.48
Temperature sum	−2.37 ± 2.13	10.62 ± 8.03		−0.19 ± 102.63	−14.00 ± 24.92
Humidity sum	−1.61 ± 1.52			0.10 ± 38.56	−12.20 ± 18.53
Rain sum	0.47 ± 0.26	−1.91 ± 5.74	0.69 ± 1.99	0.50 ± 297.64	−2.00 ± 2.16
Wind sum	0.22 ± 0.19	−0.95 ± 3.08	−0.40 ± 1.07	0.56 ± 314.03	−1.85 ± 1.38

Effect size ± SD is given for each parameter. Positive values indicate an increase in relation to the respective reference value whereas negative values indicate a decrease. All other parameters are as explained in the legend of Table 1. ***p < 0.001