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. 2016 Nov 8;6:36550. doi: 10.1038/srep36550

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This work is licensed under a Creative Commons Attribution 4.0 International License. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in the credit line; if the material is not included under the Creative Commons license, users will need to obtain permission from the license holder to reproduce the material. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/

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Samples from three plots in the same tidal flat are represented by different colors. Samples from the four seasons are represented by different shapes of symbols.