Learned tuning curves continue to exhibit characteristic LMN HD cell properties (i.e., turn-dependent offset and peak firing rates) when subject to learned spatially modulated feedback. Colors represent the different turning states (compare with Fig. 3): orange represents CW turns; green represents CCW turns; blue represents STR/straight/still. See arrows. A1, LMNcw in the circular environment (CW tuning curves offset CCWwise), A2, LMNccw in the circular environment. B1, B2, Same for the rectangular environment.