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Structure of the six nucleotide-binding sites of the proteasome. (A) Overview of six nucleotide-binding sites in the AAA-ATPase heterohexamer of the SA state. Bound nucleotides are shown in stick representation superimposed with cryo-EM densities of the nucleotide that are shown in blue mesh. (B) Close-up views of nucleotide conformations in the six nucleotide-binding sites in SA. ATP is tentatively modeled into the nucleotide density of each Rpt subunit in SA.
