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Position of the intrinsic arginine finger in crystal structures. In the active GTPγS-bound state (PDB ID code 1GIA), Arg178 is oriented away from the substrate toward a glutamate (Glu43). In the aluminium-tetrafluoride (AlF4−)-stabilized intermediate (PDB ID code 1GFI), Arg178 points toward the substrate. In the inactive GDP state (PDB ID code 1GP2), Arg178 is again flipped away from the substrate.
